Name: ___________________________
Chemistry 360, Exam 3
March 14, 1997
General instructions: Read each question carefully and give the answer in the expected format. Show your setup for all calculation problems so I can give partial credit if necessary. Include units of measurement on all answers, If you need a calculator, pencil, eraser, more paper, etc., ask. Abbreviations are discouraged to guard against misinterpretation.
Part A.
1. Draw out the Embden-Meyerhof pathway under anaerobic conditions, detailing the enzymes involved and their coenzymes, and the names and structures of the intermediates. (25 points)

2. List sequentially the enzymes involved in the tricarboxylic acid cycle, beginning with pyruvate as the substrate. (10 points)
        pyruvate dehydrogenase
        citrate synthase
        aconitase
        isocitrate dehydrogenase
        a-ketoglutarate dehydrogenase
        succinate thiokinase
        succinate dehydrogenase
        fumarase
        malate dehydrogenase

3. Schematically diagram the electron transport chain, indicating the major molecules, enzymes of each site and where ATP are produced. (10 points)

4. "Metabolism" is a general term. List the activities which comprise this term. (5 points)
    Anabolism and catabolism
    1.  Catabolism of dietary macromolecules into building block molecules
    2.  Anabolism of building block molecules into macromolecules
    3.  Synthesis of building block molecules
    4.  Synthesis and degradation of specialized molecules

5. Using a tabular format, compare and contrast fatty acid anabolism and catabolism with respect to products and reactants, cofactors, location, differing intermediates and how reactions are carried out. (10 points)
                                            Anabolism                                    Catabolism
reactants -> products    acetyl CoA -> palmitic acid        fatty acyl CoA -> acetyl CoA
cofactors                            NADPH                                FAD and NAD+
location                            cytoplasm                                    matrix
differing intermediates             D                                                L
how reactions are carried out    1 polypeptide with                 multiple enzymes
                                                    multiple enzyme
                                                    activities

6. What is the net energy yield (ATP equivalents) from a triglyceride molecule composed of one palmitic, one stearic and one oleic acid molecule. Ignore the energy contribution from glycerol. (10 points)
palmitic - 7 cycles                        stearic - 8 cycles         oleic - "8" cycles
    -2 activate                                        -2                             -2
    7x17 ATP per cycle                         8x17                         8x17
       12 ATP from other acetyl CoA         12                            12
                                                                                            -2 for double bond
    _________________                    ______                        ______
        129                                                146                            144

total 421 ATP net

7. Demonstrate the conversion of palmitic acid to stearic acid by outlining the molecules involved, and enzymes and coenzymes involved in their interconversion. (10 points)

Part B. Multiple Choice (3 points each). Circle the letter of the most correct answer.
1. Hexokinase
    a. is not regulated
    b. is regulated by NADH
    c. is regulated by glucose-6-phosphate
    d. is regulated by ATP
correct answer is C

2. An amphibolic pathway
    a. is involved in anabolism
    b. is involved in both anabolism and catabolism
    c. is involved in catabolism
    d. is involved in neither anabolism or catabolism
correct answer is B

3. The energy yield of glycolysis aerobically
    a. is 8 ATP
    b. is 6 ATP
    c. is 2 ATP
    d. is 2 ATP more than anaerobically
no correct answer - depends on transporter used

4. The chemiosmotic coupling hypothesis
    a. says that two protons cause the synthesis of an ATP
    b. says that mitochondria must be intact.
    c. has a great deal of experimental evidence to back it up
    d. a and b
    e. b and c
    f. all of the above
correct answer is F

5. ß-oxidation
    a. requires carnitine
    b. requires NADP+
    c. requires FAD
    d. requires ATP
    e. a and c
    f. all but b
    g. all but d
correct answer is F

6. Phosphofructokinase
    a. is the rate limiting step of glycolysis
    b. is inhibited by ATP
    c. is activated by NAD+
    d. all of the above
best answer is D, but we did not discuss all these molecules

7. The rate limiting step of the Krebs cycle is
    a. a-ketoglutarate dehydrogenase
    b. succinate thiokinase
    c. citrate synthase
    d. fumarase
correct answer is C

8. The Krebs cycle
    a. occurs in the cytoplasm
    b. occurs in the mitochondria
    c. occurs in endoplasmic reticulum
    d. all of the above
correct answer is B

9. One of the products of ß-oxidation of odd-chain-length fatty acids is
    a. FADH2
    b. succinyl CoA
    c. propionyl CoA
    d. a and b
    e. b and c
    f. a and c
correct answer is C

10. Fatty acid elongation occurs
    a. in mitochondria
    b. in the endoplasmic reticulum
    c. in the cytoplasm
    d. in the intermembrane space of mitochondria
correct answer is B